《园林树木学西农-吉文丽》第一章.ppt

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1、pandectChapter 1 Taxonomy and Classification of Landscape trees(2 periods)Methods of Classification:Natural classificationArtificial classification,Taxonomy(分类学):the classification of organisms in an ordered hierarchical system that indicates natural and evolutionary relationships between them.-natu

2、ral classificationClassification(分类):according to the function of woody landscape plants in urban greening.-artificial classification,As we know,there are about 8000 woody landscape plants in our country.Of these,3000 species have been cultivated.We also introduced more than 1000 cultivars from fore

3、ign country.More than 2000 species in Xishuangbanna botanical garden are collected,but only several hundreds species are applied to urban area.In order to grow more woody species in landscape architecture,we should know the names,characteristics of woody plants first.Correct plant is very important

4、for application.It is worse and more dangerous to people for using wrong plants as food.,For example,木本香料并做调料品的八角和莽草都是同科（八角科）、同属（八角属）、的常绿乔木。形态很相似（八角的聚合 果8，先端钝尖或短尖；莽草聚合 果10-13，先端是细长弯曲的尖头）。如不能正确区别，误食莽草可置人于死地。所以需正确鉴定，进行分类。,We will review the principles of systematic classification,the plant nomenclatur

5、e,and keys first,then learn how to construct and use dichotomous key(分支式检索表).分类学（taxonomy）可以定义为研究和描述有机体的变异、探讨这种变异的因果关系、并运用所掌握的资料去建立某个分类系统的学科。分类学和系统学（systematic）这两个术语现在常被不严格地作为同义词并用，如果要说两个术语有些什么不同的含义，那么分类学常用来指分类理论和实践（包括它的基础、原理、步骤和规则）而不是它的最终产物（分类系统）；系统学是有机体的种类和多样性以及它们间全部亲缘关系的研究（simpson 1961），比分类学含义多少宽

6、广些。,The principles of systematic classification(系统分类的基本原则)Systematics(系统分类学):the study of relationships of organisms and evolutionary processes.The taxonomic organization of species is hierarchical.Each species(Prunus persica桃,for example)belongs to a genus(Prunus桃属,李属,樱属,梅属),each genus belongs to a

7、 family(Rosaceae蔷薇科),each family belongs to an order(Rosales蔷薇目),each order belongs to a class(Dicotyledoneae双子叶植物纲),each class belongs to a phylum(Spermatophyta种子植物门),and each phylum belongs to a kingdom(Plantae植物界).,Concept of species物种的概念：大家所接受的概念是：种是自然界中客观存在的一种类群，这个类群中的所有个体都有着极其相似的形态特征和生理、生态特性，个

8、体之间可以自然交配产生正常的后代而使种族延续，它们在自然界中占有一定的分布区域。人们就以这种客观存在的类群称为种，也是作为分类的基本单位。Home workhttp:/sorrel.humboldt.edu/kll1/speciesdef.html,Important Plant Classification TerminologyGenus:Weakly defined as a group of plants containing one or more species.The species have more characteristics in common with each ot

9、her than they do with species of other genera in the same family.Similarity of flowers and fruits is the most widely used feature of comparison.A genus may contain a single species(e.g.,Ginkgo biloba)or more than 100(e.g.,Rosa spp.).The plural of the singular genus is genera,Species:A group of indiv

10、idual plants that are fundamentally the same,separated from other closely related species by distinct morphological differences.All individuals in a given species are not identical,but can express variation in certain traits.Species is abbreviated sp.(singular)or spp.(plural).,Variety:A population o

11、f plants within a species that display clear differences,differences that occur in natural populations.A variety name is written in lower case,italicized or underlined,and preceded by the abbreviation var.For example,Honeylocust is Gleditsia triacanthos whereas the Thornless Honeylocust is Gleditsia

12、 triacanthos var.inermis(Latin for lacking thorns).Subspecies(abbreviated ssp.)is for practical purposes equivalent to variety.,Forma:Describes sporadic variations,such as an occasional white flowered plant in a population of a normally red-flowered species.For example,Quercusglauca Thung.(青岗栎)is us

13、ually big leaves in nature,but small leaves plants occur naturally,being referred to as Quercus glauca Thung.f.gracilis Rehd.et.Wils(小叶青岗栎).,Cultivar:A term coined by Liberty Hyde Bailey and derived from the term cultivated variety.It is defined as an assemblage of cultivated plants,which are clearl

14、y distinguished by one or more characters,and which when reproduced(sexually or asexually)retains its distinguishing characteristics.A yellow stem form of Redosier Dogwood is Cornus sericea Flaviramea(金枝梾木）.The cultivar name is not italicized or underlined and has single quotes on either side of the

15、 name.Cultivar is abbreviated cv.so the plant could also be named with the single quotes deleted,such as,Cornus sericea cv.Flaviramea.,Hybrid:This is a plant that has originated from a cross(or hybridization)between two species within a genera(Interspecific Hybrid),or two different genera(Intergener

16、ic Hybrid).The small multiplication sign x used between genus and specific epithet names simply indicates that the plant is an interspecific hybrid origin(Example:Abelia x grandiflora).An intergeneric hybrid is written with a large multiplication sign X before the genus name(Example:X Cupressocypari

17、s leylandii.),1.种的学名如：银杏 Ginkgo biloba L.多为名词 多为形容词 命名人缩写2.变种的学名：在种名之后加上Var.(varietas)符号再加上变种词 如：樟子松是欧洲松的变种 Pinus sylvestris Linn.var.mongolica Litr.欧洲松 变种符号 变种加词常写为 Pinus sylvestris var.mongolica 3.变型的学名：f.(forma)小叶青岗栎 Quercus glauca Thung.f.gracilis Rehd.et.Wils.符号 变型加词,Patents and Trademarks:Pate

18、nts give exclusive rights and protection to an inventor to make,use,and sell an invention.For 20 years after the date indicated on a plant patent,only the patent holder may commercially raise or sell a patented plant.Others may do so through license or royalty agreements with the patent holder.Trade

19、marks provide another form of protection.The plant name can be trademarked and such names cannot be used as a name for any other similar plant.,Trademarked plant names are indicated by use of a trademark(),such as Betula nigra Heritage,whereas patented plants are designated with a registered tradema

20、rk().Confusing to some is the fact that patented cultivars must be given,in addition to the registered trademark name,an official cultivar name registered according to the International Code of Nomenclature for Cultivated Plants(ICNCP国际栽培植物命名法规).In the case of Betula nigra Heritage,the actual cultiv

21、ar name approved by the International Cultivar Registration Authorities(ICRAs国际品种登录权威)is actually Cully.,Plant Recognition Learning to recognize different kinds of plants in the nursery and landscape is an important skill that requires close observation and knowledge of plant parts.Landscape ornamen

22、tals vary greatly in overall shape and habit,ranging from sprawling groundcovers to mounding shrubs,to columnar to weeping trees.,Even when the general shape is similar,landscape plants can differ in numerous other traits,including the shape,color,and size of leaves,flowers,and fruits,and the color

23、and texture of bark and twigs.Notwithstanding,knowledge of these characteristics alone may not be enough to clearly separate one type of plant from another,especially when environmental or cultural factors like pruning have altered the plants natural form and growth,or the plant is very young or old

24、.,For these reasons,additional morphological features often need to be examined for accurate plant identification,like leaf arrangement and type,and leaf shape,and leaf apices,leaf bases,leaf margins,type of leaf venation,shape of leaf scars and vascular bundles,terminal and lateral bud shape,size,a

25、nd color,and flower and inflorescence morphology.,3分类系统上的等级（以桃为例）界-植物界 门-种子植物门 亚门-被子植物亚门 纲-双子叶植物纲 亚纲-离瓣花亚纲 目-蔷薇目 亚目-蔷薇科 亚科-李亚科 属-梅属 亚属-桃亚属 种-桃,Biological system of Linnaeus(Taxonomic System)In the formal nomenclature of any species,living or fossil,taxonomists follow the biological system of Linnaeu

26、s.Kingdom Phylum(phyla)Class Order Family Genus(genera)Species Subdivisions:Sub-(亚）,super（超）,Plant Nomenclature Introductory CommentsThe overall objective of this lecture is to help you learn about plant nomenclature and classification.The information presented here provides an important foundation

27、for future learning about landscape plants.You will learn principles that govern the proper writing of scientific names(i.e.Latin binomial),skills that are essential for communication within the landscape industry.You will also become familiar with the scientific basis for classifying plants into di

28、stinct groupings.Spend time putting as much of this information as possible to memory before moving on.,Binomial nomenclature(二名法，双名法）:Each species is defined by two names:the generic and specific names with Latin.In full taxonomic nomenclature the authors name and the date of the publication are gi

29、ven after the species,e.g.Squamularia grandlis Chao,1929 Generic name+Specific name+Authors name+Date Both generic and specific names in italic,Law of priority(优先律）All the nomenclature of organisms follow the law of priority:The first published name is valid,Plant NamesPlants can be described using

30、descriptors such as tree,shrub,vine,groundcover,evergreen,deciduous,annual,perennial,woody,herbaceous,hardy,tender,etc.Single word descriptions like these are useful but contribute only little to separating one plant-kind from another.,For example,many quite different plants could all be referred to

31、 as trees.More thorough descriptions using many words,as was done in early scientific naming systems,are too complex and cumbersome for common usage.Today,each plant kind is given a unique scientific name.Carolus Linnaeus(in his Species Plantarum,1753)first developed the scientific system we now use

32、 to name plants,referred to as the Linnaean Binomial System of Nomenclature.,Using this system,plants are given two Latinized names that create a binomial.In the Latin binomial,the first name is called the genus and the second is called the specific epithet.The generic name and the specific epithet

33、combine to form the species name.For example,the species name for the North American Sugar Maple is Acer saccharum.The Latin genus Acer is the ancient Latin name for maple(perhaps meaning hard as is the wood),whereas the specific epithet saccharum is latin for sugar or sugar cane.,Since few people s

34、peak Latin,an obvious question might be,why not simply use common English names,like Sugar Maple,instead of scientific Latin names?The primary problem with common names is that they are often commonly used in only localized regions.,For example,the Sugar Maple is often called Rock or Hard Maple in d

35、ifferent parts of the United States.The American Hornbeam(Carpinus caroliniana)is also called Blue Beech,Musclewood,Water Beech,or Ironwood in various regions.Dirr notes in his Manual of Woody Landscape Plants(1998)that the European White Waterlily(Nymphaea alba)has 15 different common English names

36、,44 French,105 German,and 81 Dutch names,a total of over 245 common names,To compound the problem,identical common names in different regions often refer to completely different plants.The potential for confusion in the communications between people living in different geographical regions is obviou

37、s.To standardize plant names around the world,a single species name,a Latin binomial,has been established for each plant according to the International Code of Botanical Nomenclature(ICBN)by the activity of authorities coordinated in part through the International Association for Plant Taxonomy(IAPT

38、).The registering of culivar names is the resonsibility of the International Cultivar Registration Authorities(ICRAs).,国际植物命名法规International Code of Botanical Nomenclature(ICBN)一种植物只能有一个合法的名称，其它名称都作为异名 一种植物若有几个名称，应按优先律（最早发表的为合法名称）处理。所有植物名都要用拉丁名或拉丁化的名称。给每种植物命名，必须有模式标本。合法名称的成立必须在正式国家刊物、杂志、著作发表，必须有拉丁文记

39、载。,中名的命名原则 一种植物只应有一个全国通用的中文名称，至于全国各地的地方名，可任其存在而称为地方名。中文属名是植物中名的核心，在拟定时选择使用广泛、形象生动与形态、生态、用途有联系的中名作属名。名称中尽量避免迷信色彩及纪念古人或今人的名称,目前还存在混乱。,Writing a Species NameThe species name(i.e.Latin binomial)is italicized or underlined.The first letter of the genus is in upper case whereas the specific epithet is in

40、all lower case letters.As an example,the species name for Sugar Maple should be written as:Acer saccharum or Acer saccharumA name often following the Latin binomial,as in Acer saccharum Marsh.,is the abbreviated name of the individual who first assigned a scientific name to the plant,in this case Hu

41、mphrey Marshall.The letter L.in the Latin name Hedera helix L.identifies Linnaeus as the authority.The authority name is not typically included in nursery and landscape literature.,In the seed plants,there are many systems of plant taxonomy,like A.Engler;J.Hutchinson;A.Takhtajan;ACronquist;Zheng Wan

42、jun,etc.Most common systems of plant taxonomy are A.Engler;J.Hutchinson 在种子植物门中，有许多分类系统。象哈钦松分类系统、恩格勒分类系统、塔赫他间分类系统、克朗奎斯特系统、郑万钧系统等。目前最常用的是哈钦松分类系统和恩格勒分类系统。,自然分类系统中两个主要系统的特点简介 1.恩格勒（Enger）系统 德国的恩格勒编写了两本巨著植物自然分科志和植物分科志要系统描述了全世界的植物，内容丰富并有插图，很多国家采用了这个系统。特点如下：在被子植物中，单性而无花被的为原始特征，所以将木麻黄科、杨柳科、桦木科等放在木兰科、毛茛科之前。

43、认为单子叶植物比双子叶植物原始。1964年改变，把双子叶植物放在前边，便于同其他植物学家统一。目与科的范围较大。该系统较稳定而实用，所以在世界各国及中国北方多采用，如：中国树木分类中国植物志中国高等植物图鉴等书均采用本系统。,2 哈钦松（J.Hutchinson）系统 英国的哈钦松在其著作有花植物志科中公布了这个系统。特点如下：认为单子叶植物比较进化，故排在双子叶植物之后。在双子叶植物中，将木本与草本分开，并认为乔木为原始性状，草本为进化性状。认为花的各部呈离生状态、花的各部呈螺旋状排列、具有多数离生雄蕊、两性花等性状均较原始；而花的各部分呈合生或附生、花部呈轮状排列、具有少数合生雄蕊、单性花

44、等性状属于较进化的性状。,哈钦松（J.Hutchinson）系统 认为在具有萼片和花瓣的植物中，如果它的雄蕊和雌蕊在解剖上属于原始性状时，则比无萼片与花瓣的植物为原始，例如杨柳科等的无花被特征是属于退化的现象。单叶和叶呈互生排列现象属于原始性状，复叶或叶呈对生或轮生排列现象属于较进化的现象。目与科的范围较小。目前认为该系统较为合理，但原书中没有包括裸子植物。中国南方学者采用的多。如：南方植物园、广州植物志园林树木1000种、树木学、海南植物志等都是哈钦松分类系统。,The system of Engler and Prantl has servedly commanded universal

45、attention.Their system has been widely accepted,though in recent years it has been subjected to some criticism.It is generally admitted that many anomalies in the older arrangements were adjusted and that great contributions were made to our knowledge of Taxonomic Botany,Mr.Hutchinson has ventured t

46、o express his dissent from Engler and Prantls views in this book and has returned,wisely as I think,to the position adopted by the authors of the Genera Plantarum,since he regards the Ranales on the one hand and the Magnoliales on the other as the starting-points of his phylogenetic arrangement,Iden

47、tification is the recognition of characteristics of organisms and the application of a name to an organism with those particular characters The dichotomous key is the method employed for identifying unknown organisms.The evolution of keys has been the result of work by taxonomists who study the char

48、acteristics of organisms at some taxonomic level(=category)and often develop keys for their identification.,This exercise introduces you to the construction and use of a dichotomous key.It is not meant to provide students a method of quick memorization of scientific names.The proficiency level of us

49、ing keys,and learning names,is a reflection of the amount of time one spends keying organisms.The more time spent using keys,the more familiar the characteristics of organisms become,and the more proficient one becomes with keys.,植物分类检索表 定距检索表 本检索表中，对某一种性状的描述是从书页左边一定距离处开始，而与其相对的性状描述亦是从书页左边同一距离处开始；其下

50、一级的两个相对性状的描述又均在更大一些的距离上开始，如此逐渐下去，距书页左方愈来愈远，直至检索出所需要的名称为止。,A1植物体无根、茎、叶的分化-低等植物门 B1植物体不为藻类和菌类所组成的共生体 C1植物体内有叶绿素或其它光合作用-藻类植物门 C2植物体内无叶绿素或其它光合作用-菌类植物门 B2植物体为藻类和菌类所组成的共生体-地衣植物门A2植物体有根、茎、叶的分化-高等植物 B1植物体有茎、叶，而无真根-苔藓植物门 B2植物体有茎、叶，也有真根 C1不产生种子，用孢子繁殖-蕨类植物门 C2产生种子-种子植物门,平行检索表：本检索表中每一相对性状的描写紧紧并列以便比较，在一种性状描述之后即列